MP3 vs FLAC: Lossy VS Lossless
Not all speakers can reproduce sounds accurately enough to explicit difference between FLAC and MP3, and not everyone knows what to search for in a song to see if it was compressed or not.
If you just want to hear a song in a general way, ie words, rhythm, general feel, you probably won't notice any difference.
If you want to hear the clear cymbal sound, completely independently from the other sounds, if you wanted to isolate the bass guitar and separate it from bass drums instances and background static, or in a general way, hear every note "alone", independently, you are going to need FLAC(or wma lossless) and good speakers/cans.
Edit: Also, a note I forgot, if you are listening to 320kbps .mp3 files, it's going to be close to impossible to distinguish their sounds from FLAC, unless you had audiophile grade equipment.
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