PCIe 4.0, Ram, GPU, and MOBO slots.
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Solved by Chris Pratt,
RAM is not over PCIe. You also don't generally need to choose between the graphics card and SSDs in terms of PCIe 4.0. Assuming the board supports PCIe 4.0 at all (you didn't specify AMD or Intel), it's either going to be all PCIe 4.0, or at least PCIe 4.0 on the CPU lanes, which gives you at least one x16 slot and an M.2. If the chipset lanes are also PCIe 4.0, then there's no limitations.
That said, even PCIe 4.0 graphics cards don't really saturate over PCIe 3.0, and right now at least, there's very little point to PCIe 4.0 SSDs, unless you're doing workloads that are very I/O heavy, like editing 4K+ raw video.
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