Reassurance
ASUS MAXIMUS VII HERO
This is what it says on their specifications site.
*1: The PCIe 2.0 x16 slot (PCIEX4_3) shares bandwith with PCIe 2.0 x1 and M.2 slot. The default setting is Auto Mode, which automatically optimizes the system bandwidth. If you install a PCIe 2.0 x4 device, the system will automatically detect and disable PCIe 2.0 x1 and M.2 slot.
*2: The PCIe 2.0 x1 slots (PCIEX1_1/2/3) will be disabled when PCIe 2.0 x16 slot (PCIEX4_3 ) operates under x4 speed or M.2 mode.
*3: The supported functions depends on the CPU installed.
*4: These SATA ports are for data hard drivers only. ATAPI devices are not supported.
*5: 1 x USB2.0 ports at mid-board shares with ROG extension (ROG_EXT) port.
So if you put the graphics cards in the first two PCIe x16 slot, they will both run at x8 speeds.
Only if you put it in the third PCIe x16 slot (the black one), which is 2.0, it will "optimize system bandwidth", also, if you use that slot, it'll disable the PCIe x1 slots, but you won't be using it so it does not affect you.
3 is almost irrelevant since you are using a 4790K. The other points do not matter regarding PCIe configurations.
It will run your setup perfectly.

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