NAS not booting with HDD, only without HDD
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Solved by Undergrid,
Under linux (and IIRC, it's been a while) USB drives generally show up as /dev/sd(x) devices. I would guess that adding the hard drive has changed the location of your boot USB from /dev/sda to /dev/sdb (the screen is certainly showing something on sdb) and the boot sequence can't find the next thing it needs because it's looking for it on the HDD.
Check your BIOS with both the HDD and USB drive plugged in, see if there's anything that will allow you to change the boot order and/or drive order (different BIOS's call it different things).
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